Zoanthids can and will readily propagate on any surface, glass, rock, skeletal sps, powerheads, over flow boxes etc, and even substrate as seen in the picture below by TCU. I had polyps which grew up the glass and almost out of my tank several years ago.
I have actually witnessed them growing faster on the substrate. In my opinion, the surface doesn't dictate the growth pattern or rate of growth. The tank's maturity, lighting, current and parameters can and will determine every polyps rate of growth in captivity.
If your goal is complete coverage of your substrate, once they begin to spread, do not touch, cut or frag them, just let them grow. Stressing a completely happy and thriving colony
CAN stunt it's growth in a heartbeat.
Good Luck
Mucho Reef