I swear I am not a stockholder in marinebreeder.org, but check out their site...think my last 4 post have all linked to them.
anyway.....does the male have a calm dark(ish) spot to hang out in? Lots of light on your 125, maybe he feels exposed constantly? Our male is usually found in the darkest lowest traffic area during holding.
Maybe he isn't getting fat enough between holdings? When I was fighting algae through major food witholding, the male seemed to do what yours is doing...except maybe holding for a week prior to swallowing.
Overly active nighttime tankmates apparently cause the male to stress....certain tangs especially I have read. Tang being a ***** all night to the only other fish swimming around may be stressful for the male....I see a tang for your avatar....it looks big, and seems to be looking for someone to harass in 'his' tank. good luck...no one right answer.