Light doesn't seem to have much to do with it. However, darkening is often linked to anemone association (even dependent upon anemone species) as well as age. Both factors seem to be primary contributors to darkening.
My clowns have had tans before. When the anemone moves under the lights the become black on the top. When the anemone moves to more shaded areas they return to thier bright orange.
My old female (Requiescat In Pace) was always bright orange until I added my anemone. Now here's another: when my female died, my male became even darker.
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