The factors that effect PE remain something of a mystery along with coral coloration. the real answer is we don't know though you will hear and read many oversimplified explanations. We don't even know if it is a sign of coral health or possibly even a sign of distress. It probably is the result of the interaction of many variables including flow, water chemistry, lighting, tank inhabitants, nutrient density, etc., etc. Nate made a very interesting observation with which I concur; that the presence of angelfish decrease PE.