hi there.
just thought id throw in my 2 cents for what its worth with my recent experience with bleaching and browning.
firstly while in fiji on one of the smaller islands there coral reef were bleaching and as it turned out this was due to silt coming from down river making the water murky so that light was unable to reach the coral effectively and this lack of light was bleaching the reef. perhaps the zooxanthellae was being expelled under the stress which lead to bleaching which lead to death
on my 4ft ive been running 4x 54w t5s and have had great growth in my sps and other corals and just the other day i was playing around with some metal halides i had lying around and decided to stick em over me tank to see how they looked
i had them running for 2 hours and silly me i didnt put the glass plate diffuser on the halide which i believe filters out the harmful uv spectrum
to my shock the next day my sps had partially turned brown and many other corals had been effected quite dramatically.
basically they got a sunburn much like we do if we spend too much time at the beach ... well an aussie beach.
from this i can assume that too much light seemed to drive out the zooxanthellae turning the coral brown
and too little light as in the fiji reef also leads to bleaching but do not know weather corals browned before bleaching in that case so no real conclusion can be formed due to too many unknown variables like water quality ect.
i also beleive as the tempratures are heating up in the oceans this is also bleaching corals on many reefs globally
the only solid conclusion i seem to be able to come to is that corals under stress for extended periods of time weather too much or too little light or poor water conditions or extreme temperatures will experience both browning and bleaching resulting in death.
personally i think we know too little about these animals to make absolute conclusions
just my thoughts but i could be off and always willing to listen to others experiences.
Typical response would be to bring in more zooanthelle to address the limited light.
It's more likely high temperatures, other industrial/agricultural pollutants or even sunscreen (yes, it's a real problem in tourists areas) were the cause of bleaching, not suspended silt.