Looks like flow to me. Bet if you gave them less flow, the skirts would shrink right up tight. Looks like they could appreciate a little more light too. Bet they will color up better if you do.
It got way worse to where they weren't even opening anymore but something happened and they are beginning to recover. I am not exactly sure what caused them to make a come back. I am kind of thinking that because I had a Flower anemone and Maxi Mini Anemone fighting with each other and they have since stopped and the zoas are getting better.
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