Hi Matt,
I may be very wrong here. If so, please correct me. Nitrifying bacteria need n, p and organic carbon (in addition to potassium, iron and some other elements) to live and thrive. Bacteria may be able to get their carbon needs from CO2 or even alkalinity (hence drop in alkalinity due to nitrogen cycle), but usually need some organic carbon source to multiply rapidly.
In my opinion, there are two possibilities here. One is that as you said with elevated n and p, you need more space for bacteria to colonise to reduce these nutrients. This assumes that your system is not carbon limited. If this assumption right, then you should indeed see drop in nutrients as soon as colonisation of nitrifying bacteria takes place on (or in?) the new media. The other possibility is that your system is carbon limited in which case adding more media for nitrifying bacteria will not make much different because without additional organic carbon bacteria population will not increase to consume n and p.
If n or p, or both are in short supply (e.g. zero), then adding organic carbon will not help either because bacteria cannot multiply without sufficient n and p in the water column.
Most reef set up are carbon limited with plenty of n and p. This is why I often assume that the second possibility is more likely than the first. Again, I may be wrong in this assumption.
Cheers
Bülent
I agree with parts of this, Bulent but not all..
And before I begin, please disregard the facile explanations if you are already aware.. I'm just trying to be as explanatory as possible.. No offense! I am not a biologist, this is only my understanding of how it works..
First, let's make sure we are talking about the right thing, here. It's denitrifying bacteria we are talking about, not nitrifying bacteria.
Nitrifying bacteria, that live mostly on the surfaces of everything in the aquarium and that require lots of oxygen to breath are the bacteria that convert ammonia to nitrite and then nitrate. Denitrifying bacteria, that grow in the very low oxygen areas of the tank (like in rocks, dsbs and other media like matrix or siporax) and use oxygen from no3 (nitrate) to breath are what we are talking about. It is the denitrifying bacteria colonizing the low oxygen areas and pulling the oxygen molecule out of the nitrate and phosphate that reduce the nutrients. They seem to like the oxygen molecule in the no3 better than the po4.. Not sure why.. Maybe on a chemical level it is easier to extract o from no3 than po4.. (that's beyond me..)
The following is my understanding not necessarily fact.. When talking about a captive reef with elevated or increasing nutrients (no3, po4) , it is almost impossible to have a carbon limited system. The very nature of a captive system with many organisms being fed regularly is such that some carbon is always present- especially if nutrients are elevated (fish foods and coral foods are also carbon sources). The available carbon may not be the best type of carbon like simple carbohydrates that come from sugar/vodka/pellets/vinegar etc but there is always enough to allow denitrifying bacteria to colonize all available space.
WITHOUT an actively added carbon source, the bacteria will sort of find a balance between available space to live given the amount of n and p there is to consume. As n and p climb, the bacteria can only colonize the available space and then reach capacity. If there is no more space, their numbers stop increasing. And if there is no more n or p, the same will occur. But given more space and available no3 and po4, the bacteria will continue to spread and increase in numbers. And thus be more capable of reducing the nutrients.
The addition of an external carbon source, either allows the existing denitrifying bacteria to go into overdrive OR allows different bacteria to thrive and as they populate the tank (sometimes clouding the water or producing bacterial slime in the shaded areas), they consume even more n and p.
To my mind, a 'carbon limited' system is a system with no waste producing animals in it... Not a reef tank.. In other words, for our intents and purposes, not possible..
If a system is phosphate or iron limited (or anything that will not allow bacteria to flourish) then yes adding more matrix or other surface area for denitrifying bacteria is useless.. Even adding a carbon source will be useless..
So, yes, bacteria need some po4, no3, carbon, iron.. Etc to thrive and without one of these, the bacteria stop working but I believe that carbon is almost always available.
I guess that where we aren't on the same page is with the concept of 'carbon limited'. I don't see that as a possibility in a full blown reef system.
So, I feel that as long as there is extra surface area and available n and p, bacteria will colonize that surface area and reduce the n and p.. Given enough surface area, denitrifying bacteria will reduce all available nitrate to 0 - as well as pull p down, but usually not as efficiently as n.
I guess where we differ is that I don't see carbon ever being a limiting factor whereas you feel that it is a common limiting factor..