I'll try to answer this... kinda scary, since I'm just been asking questions...
It has to do with linear flow. The more linear flow the better. So, let's say your oveflow is coast to coast on a 48" tank, and you have teeth that are 1/4" and a 1/4" apart, you have reduced the linear flow to 24". A 24" sheet of water also going to be taller as it runs over the overflow, where on the same tank it would be a much thinner sheet of water if the linear flow was 48". This is suppose to be a much more efficient overflow (for reasons I am unable to explain at the moment).
So, what I've seen in some of the posts, is that people will use eggcrate to help keep critters out of the oveflow... so rather than having 1/4" teeth, you only have say 1/16" teeth...
Hope that helps.